TABLE OF CONTENTS

• 1. Introduction
  
• 2. The metaphor of adaptive landscapes
  
• 3. Steps towards speciation on rugged adaptive landscapes
  
  • Stochastic transitions between isolated adaptive peaks
    
  • Shifting-balance theory scenario
    
  • Founder effect speciation scenario
    
  • Peak shifts by selection
    
  • Divergence driven by selection for local adaptation
    
• 4. The Bateson-Dobzhansky-Muller (BDM) model
  • Allopatric speciation in the BDM model
    
  • Parapatric speciation in the BDM model
    
• 5. Holey adaptive landscapes
  
• 6. Speciation on holey adaptive landscapes
  
  • Multiallele versions of the BDM model
    
  • Multilocus versions of the BDM model
    
• 7. Concluding comments
  
• Box 1: Fixation of an underdominant mutation
  
• Box 2: Peak shift in a quantitative character
  
• Box 3: Terminology
  
• Box 4: Russian roulette model